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Spinosaurus (meaning 'spine lizard') was a theropod dinosaur genus that lived in what is now Egypt, from the Albian to early Cenomanian stages of the Cretaceous Period, about 95 to 93 million years ago. According to a study by dal Sasso et al. (2006), it was the largest of all carnivorous dinosaurs by a significant margin, even larger than Tyrannosaurus rex and Giganotosaurus. Spinosaurus was the longest theropod, measuring 15 to 17.4 metres (49.2 to 57.1 feet) long and standing at 5 to 6 metres (16 to 20 feet) in height. Spinosaurus reached weights up to 9 tons. The distinctive spines of these animals (large bones extending from the vertebrae) grew up to 2 m (6.6ft) long and were likely to have had skin stretching between them, forming a sail-like structure, although some have suggested they were covered in muscle and formed a hump or ridge. Spinosaurus provides the name of a family of dinosaurs, the Spinosauridae, of which other members include Angaturama (probably synonymous with Irritator), Baryonyx, Irritator, Suchomimus, and Siamosaurus.
Description Although Spinosaurus is well-known to dinosaur enthusiasts due to its unusual features, it is mostly known from remains that have been destroyed, aside from a few more recently discovered teeth and skull elements. Jaw and skull material published in 2006 show that it had one of the longest skulls of any carnivorous dinosaur, estimated by dal Sasso et al. at about 175 cm (5.7 ft). Originally found in the Bahariya Valley of Egypt in 1912, it was named by German paleontologist Ernst Stromer in 1915. Some of the fossils were damaged during transport back to the Deutsches Museum, Munich, Germany and the remaining bones were completely lost due to Allied bombing in 1944. Aside from its 'sail', notable characteristics of Spinosaurus include: Much of this is speculation based on Baryonyx and other spinosaurids, as no limb material has ever been attributed to Spinosaurus itself. Since its discovery, Spinosaurus has held the record for longest, and possibly largest, theropod dinosaur (though this fact did not reach the public consciousness until its depiction in the film Jurassic Park III and the description of a new specimen in 2006). Both Huene (1926) and Glut (1982) listed it as the most massive theropod in their surveys, at upwards of 6 tons in weight and 15 meters (50 feet) in length. Paul (1988) also listed it as the longest theropod at 15 meters (50 feet), but gave a lower mass estimate. The most recent estimates, based on new specimens described by dal Sasso et al. (2006), list Spinosaurus at 16 - 18 metres (53.3 to 60 feet) long and 7.5 - 9 tons in weight. At least one survey, as-yet unpublished, suggest that Spinosaurus reached sizes of 12 - 19 tons in weight. These high-end weight estimates are based on the fact that the vertebrae of Spinosaurus are unusually massive compared to theropods of comparable size (implying an extremely large overall mass), and that the holotype specimens are apparently sub-adult. * Diet It is unclear whether Spinosaurus was primarily a cursorial predator or a fisher, as indicated by its elongated jaws, conical teeth and raised nostrils. The only direct evidence for spinosaur diet comes from related European and South American species. Baryonyx was found with both fish scales and bones from juvenile Iguanodon in its stomach, while a tooth embedded in a South American pterosaur bone suggests that spinosaurs occasionally preyed on these flying archosaurs. Spinosaurus was likely to have been a more generalized and opportunistic predator, possibly a Cretaceous equivalent of large grizzly bears, being biased toward fishing, though it undoubtedly scavenged and took many kinds of small-to-medium-sized prey (Paul, 1988). Sail Spinosaurus sails were unusual, although other dinosaurs of the same time and area, namely the ornithopod Ouranosaurus and the sauropod Rebbachisaurus, developed a similar structural adaptation of their dorsal vertebrae. The sail is possibly analogous (not homologous) to that of the Permian mammal-like reptile, Dimetrodon, which lived before the dinosaurs even appeared (these similarities are presumably due to parallel evolution). The purpose of these sails is uncertain; scientists have proposed several hypotheses: Finally, since things in nature rarely develop for a singular reason, it is quite possible that the sail combined all these functions, acting normally as a heat regulator, becoming a courting aid during the mating season, being used to cool itself and, on occasions, turning into an intimidating device when an animal was feeling threathened. Conjecture may even allow that the sail may have changed colour, during any of these functions. Species and Specimens Two species of Spinosaurus have been named: Spinosaurus aegyptiacus ("Egyptian spine lizard") and Spinosaurus marocannus ("Moroccan spine lizard"). S. marocannus was originally described by Russell as a new species based on the length of its neck vertebrae. However, several later authors considered the length of the neck vertebrae to be variable from individual to individual and therefore consider S. marocannus to be a synonym of S. aegyptiacus *. Five partial specimens of Spinosaurus have been found, the first having been destroyed during World War II (luckily, detailed drawings and descriptions of the specimen remain). The probable size of these individual spinosaurs can be estimated using comparison to known material from other spinosaurid dinosaurs. Rauhut (2003) suggested that Stromer's Spinosaurus holotype was a chimera, consisting of dorsal vertebrae from a carcharodontosaurid similar to Acrocanthosaurus and a dentary from a large theropod similar to Baryonyx. This analysis, however, was rejected by dal Sasso and most other researchers. Spinosaurus in Jurassic Park III
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